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Some species, such as Gryllus assimilis , take off, fly, and land efficiently and well, while other species are clumsy fliers.
In other species, they may be pulled off and consumed by the cricket itself or by another individual, probably providing a nutritional boost.
Gryllus firmus exhibits wing polymorphism ; some individuals have fully functional, long hind wings and others have short wings and cannot fly.
The short-winged females have smaller flight muscles, greater ovarian development, and produce more eggs, so the polymorphism adapts the cricket for either dispersal or reproduction.
In some long-winged individuals, the flight muscles deteriorate during adulthood and the insect's reproductive capabilities improve. Captive crickets are omnivorous ; when deprived of their natural diet, they accept a wide range of organic foodstuffs.
Some species are completely herbivorous , feeding on flowers, fruit, and leaves, with ground-based species consuming seedlings, grasses, pieces of leaf, and the shoots of young plants.
Others are more predatory and include in their diet invertebrate eggs, larvae, pupae, moulting insects, scale insects , and aphids. Crickets have relatively powerful jaws, and several species have been known to bite humans.
Male crickets establish their dominance over each other by aggression. They start by lashing each other with their antennae and flaring their mandibles.
Unless one retreats at this stage, they resort to grappling, at the same time each emitting calls that are quite unlike those uttered in other circumstances.
When one achieves dominance, it sings loudly, while the loser remains silent. Females are generally attracted to males by their calls, though in nonstridulatory species, some other mechanism must be involved.
After the pair has made antennal contact, a courtship period may occur during which the character of the call changes. The female mounts the male and a single spermatophore is transferred to the external genitalia of the female.
Sperm flows from this into the female's oviduct over a period of a few minutes or up to an hour, depending on species.
After copulation , the female may remove or eat the spermatophore; males may attempt to prevent this with various ritualised behaviours.
The female may mate on several occasions with different males. Most crickets lay their eggs in the soil or inside the stems of plants, and to do this, female crickets have a long, needle-like or sabre-like egg-laying organ called an ovipositor.
Some ground-dwelling species have dispensed with this, either depositing their eggs in an underground chamber or pushing them into the wall of a burrow.
Crickets are hemimetabolic insects, whose lifecycle consists of an egg stage, a larval or nymph stage that increasingly resembles the adult form as the nymph grows, and an adult stage.
The egg hatches into a nymph about the size of a fruit fly. This passes through about 10 larval stages, and with each successive moult , it becomes more like an adult.
After the final moult, the genitalia and wings are fully developed, but a period of maturation is needed before the cricket is ready to breed.
Some species of cricket are polyandrous. In Gryllus bimaculatus , the females select and mate with multiple viable sperm donors, preferring novel mates.
Fertilization bias depends on the control of sperm transport to the sperm storage organs. The inhibition of sperm storage by female crickets can act as a form of cryptic female choice to avoid the severe negative effects of inbreeding.
Crickets have many natural enemies and are subject to various pathogens and parasites. They are eaten by large numbers of vertebrate and invertebrate predators and their hard parts are often found during the examination of animal intestines.
The entomopathogenic fungus Metarhizium anisopliae attacks and kills crickets and has been used as the basis of control in pest populations.
The diseases may spread more rapidly if the crickets become cannibalistic and eat the corpses. Red parasitic mites sometimes attach themselves to the dorsal region of crickets and may greatly affect them.
Other wasps in the family Scelionidae are egg parasitoids, seeking out batches of eggs laid by crickets in plant tissues in which to insert their eggs.
The fly Ormia ochracea has very acute hearing and targets calling male crickets. It locates its prey by ear and then lays its eggs nearby.
The developing larvae burrow inside any crickets with which they come in contact and in the course of a week or so, devour what remains of the host before pupating.
A trade-off exists for the male between attracting females and being parasitized. The phylogenetic relationships of the Gryllidae, summarized by Darryl Gwynne in from his own work using mainly anatomical characteristics and that of earlier authors, [a] are shown in the following cladogram , with the Orthoptera divided into two main groups, Ensifera crickets sensu lato and Caelifera grasshoppers.
Fossil Ensifera are found from the late Carboniferous period Mya onwards,   and the true crickets, Gryllidae, from the Triassic period to Mya.
Cladogram after Gwynne, . Schizodactylidae splay-footed crickets. Tettigonioidea katydids, bush crickets, weta. London: Aurum Press Ltd. Bowen, Rowland Cricket: A History of its Growth and Development.
Goldstein, Dan The Rough Guide to English Football — London: Rough Guides. Harte, Chris A History of Australian Cricket. London: Andre Deutsch.
Haygarth, Arthur London: Frederick Lillywhite. Major, John More Than A Game. London: HarperCollins. McCann, Tim Sussex Cricket in the Eighteenth Century.
Lewes: Sussex Record Society. Nyren, John [First published ]. Ashley Mote ed. The Cricketers of my Time. London: Robson Books. Underdown, David Start of Play.
London: Allen Lane. Webber, Roy The Phoenix History of Cricket. London: Phoenix House Ltd. Williams, Charles Cricket at Wikipedia's sister projects.
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